In Karakul and Tzurkana breeds, gene R causes grey colour and is lethal when homozygous (Serra, 1948). Nel and Louw (1953) reported a few cases of homozygous grey Karakuls which survived to maturity.

G. Ricordeau

Proceedings of the World Congress on Genetics Applied to Livestock Production, Volume 6. Round tables, , 454–461, 1982
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